生活型

herb

形態特徵

• Perennial herbs; stems erect or ascending.
• Leaves mostly unequally paired, thinly succulent, with thick cuticle on lower surface, usually ovate to lanceolate, sometimes elliptic or oblong, rarely obovate, 2-16.5 cm long, 1.2-5.4 cm wide, apex mostly caudate to long caudate, sometimes acute or cuspidate, margins entire, base obtuse to shallowly cordate, 3-nerved; petioles 0.3-7.5 cm long; stipules connate at base, broadly triangular, veinless.
• Inflorescences usually panicles of glomerules; male inflorescences 10-20 cm long, peduncles 5-11 cm long; female inflorescences 4-13.5 cm long, peduncles 2-6.5 cm long; mixed inflorescences present. Male flowers slightly compressed laterally, obovoid; sepals 4, connate about halfway, valvate, oblong- to obovate-cymbiform, corniculate. Female flowers oblong; sepals 3 or 4, connate at base, unequal, cymbiform; staminodes 3 or 4.
• Achenes slightly warty, laterally compressed, ovoid to ovoid-ellipsoid.

物候

Flowering and fruiting June to November

分布

• 台灣 : ravines, moist, shady forest floors, sunny roadsides and rocks at low to medium altitudes throughout the island.
• 世界 : Vietnam, Hainan, S. C. mainland China and Japan?.

引證標本

ILAN: Nanhutashan, Sasaki s. n. 1922; Taipingshan, Huang 3960. MIAOLI: Talu Logging Trail, Wang & Yang 4848. TAICHUNG: Lishan, Shih 2882, 2889. NANTOU: Meichi, Shimada s. n. 1918; Wushe, Kanehira & Sasaki s. n. 1918, Hsu 11387. CHIAYI: Funkiko, Shih 2964, Shih et al. 004915. KAOHSIUNG: Meishanko, Shih 2805; Shanping, Hsieh 1391. PINGTUNG: Kangkou, Kobayashi s. n. 1907; Paiwan, Matuda s. n. 1917. TAITUNG: Chihpen Logging Trail, Shih 3318; Lichia Logging Trail, Shih 3319. HUALIEN: Kuanyuan, Shih 3037; Sakatang Logging Trail, Shih 3056.

參考文獻

• YANG, YUEN-PO, SHIH, BING-LING and LIU, HO-YIH, Flora of Taiwan, second edition 2: 247. 1996.  (Fl. Taiwan 2nd edit., 台灣植物誌第二版)